Pollen is a fine to coarse powdery substance comprising pollen grains, which are the male microgametophytes of seed plants. Each pollen grain typically consists of a protective outer layer called the exine, and a single-celled male gametophyte inside, containing the generative cell and the tube cell. The exine is composed of sporopollenin, a resilient biopolymer that protects the genetic material within the pollen grain from environmental damage.
Pollen is formed within the anthers of the flower's stamen. The process of pollen formation, or microsporogenesis, begins with the development of microspore mother cells within the anther. These mother cells undergo meiosis to produce haploid microspores, which then develop into mature pollen grains through a process known as microgametogenesis.
The primary function of pollen is to facilitate the transfer of male gametes (sperm cells) from the anther of one flower to the stigma of another, ultimately leading to fertilization. This process, called pollination, can occur through various mechanisms, including wind, water, insects, birds, and other animals, depending on the plant species. The unique morphology of pollen grains, often characterized by specific surface features and ornamentation, plays a crucial role in the adaptation of plants to their respective pollination vectors.